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Price: $172.95

Hardback 599 pp.
9" x 11"



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Imprint: Sinauer Associates is an imprint of Oxford University Press

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Fourth Edition

Douglas J. Futuyma and Mark Kirkpatrick

Series : Sinauer

Extensively rewritten and reorganized, this new edition of Evolution offers additional expertise in evolutionary genetics and genomics, the fastest-developing area of evolutionary biology. Suited for an undergraduate audience, the text emphasizes the interplay between theory and empirical tests of hypotheses, helping to familiarize students with the process of science.

* Chapters marked by an asterisk have been entirely rewritten
Unit One: An Idea That Changed the World
1. Evolutionary Biology
2. The Tree of Life
3. Natural Selection and Adaptation
Unit Two: How Evolution Works
4. Mutation and Variation *
5. The Genetical Theory of Natural Selection *
6. Phenotypic Evolution *
7. Genetic Drift: Evolution at Random *
8. Evolution in Space NEW
9. Species and Speciation *
Unit Three: Products of Evolution: What Natural Selection Has Wrought
10. All About Sex *
11. How to Be Fit
12. Cooperation and Conflict *
13. Interactions among Species
14. The Evolution of Genes and Genomes *
15. Evolution and Development
Unit Four: Macroevolution and the History of Life
16. Phylogeny: The Unity and Diversity of Life
17. The History of Life
18. The Geography of Evolution
19. The Evolution of Biological Diversity
20. Macroevolution: Evolution above the Species Level
Unit Five: Evolution and Homo sapiens
21. The Evolutionary Story of Homo sapiens NEW
22. Evolution and Society
Appendix: A Statistics Primer NEW
Literature Cited
Illustration and Photo Credits

Instructor Resources:
Ancillary Resource Centre (ARC):
Image Bank:
- All figures, tables, and photos from the text
PowerPoint Slides:
For each chapter:
- All of the figures and tables, including titles and full captions
- Answers to the textbook end-of-chapter problems and discussion topics
Quiz Questions:
- From the Companion Website
Data Analysis and Simulation Exercises:
- From the Companion Website, with answers
Online Quizzes:
- Companion Website quizzes can be assigned or released for student self-study, at the instructor's discretion
- Instructors can add their own questions to create custom quizzes
Student Resources:
Companion Website:
- Chapter outlines and summaries
- Data Analysis Exercises: Inquiry-based exercises involving working with data and analyzing methods and conclusions from published papers
- Simulation Exercises: Interactive modules exploring many of the processes of evolution, and questions based on the results they
- Online quizzes
- Flashcards
- Glossary
E-book ISBN 9781605357003

Douglas J. Futuyma is Distinguished Professor in the Department of Ecology and Evolution at the State University of New York at Stony Brook. He received his B.S. from Cornell University and his Ph. D. in Zoology at the University of Michigan with Lawrence Slobodkin. Dr. Futuyma is the author of three previous editions of Evolution, as well as three editions of its predecessor, Evolutionary Biology. He received the 1997 Sewall Wright Award of the American Society of Naturalists and the 2012 Joseph Leidy Award of the Academy of Natural Sciences of Drexel University (Philadelphia). Dr. Futuyma has served as President of the Society for the Study of Evolution, the American Society of Naturalists, and the American Institute of Biological Sciences, and was elected a Fellow of the American Academy of Arts and Sciences in 1996 and the National Academy of Sciences in 2006. He has served as Editor of Evolution and is currently Editor of the Annual Review of Ecology, Evolution, and Systematics. In 2013, he was recognized as Honorary Doctor by the National University of Mongolia. An avid naturalist, his major research interests include evolution of interactions among insects and plants, speciation, and evolution of community structure.

Mark Kirkpatrick is the Painter Centennial Professor of Genetics in the Department of Integrative Biology at the University of Texas at Austin. He received his B.A. in Biology from Harvard in 1978 and his Ph.D. in Zoology from the University of Washington with Monty Slatkin in 1983. Dr. Kirkpatrick has received a Guggenheim Fellowship (1997) and a Poste Rouge Fellowship (France, 1997). He is a Fellow of the American Academy of Arts and Sciences (2008), and of the American Association for the Advancement of Science (2016). Dr. Kirkpatrick received the Sewall Wright Award from the American Society of Naturalists (2014). He has served as Associate Editor of The American Naturalist, Theoretical Population Biology, and Genetics, and on the Editorial Boards of The Annual Review of Ecology, Evolution, and Systematics and Proceedings of the Royal Society of London. Dr. Kirkpatrick's research interests are in evolutionary genetics. He has worked on sexual selection, quantitative genetics, speciation, and species ranges. Current research topics include the evolution of sex determination and chromosome rearrangements.

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Special Features

  • Addresses major themes - including the history of evolution, evolutionary processes, adaptation, and evolution as an explanatory framework - at levels of biological organization ranging from genomes to ecological communities.
  • Emphasizes the process of science to show students what science is, how it works, and why it is the most reliable way of knowing what has yet been developed.
New to this Edition
  • A stronger focus on human evolution is provided in a new chapter on the topic (Ch. 21) as well as through new examples throughout the text.
  • New and fully rewritten chapters includes the entirety of Part Two: How Evolution Works, as well as chapters on sexual selection and sexual reproduction (Ch. 10), the evolution of genes and genomes (Ch. 14), evolution and development (Ch. 15), and more.
  • A new appendix, A Statistics Primer, introduces the concept of a probability distribution, reviews how statistics are used to describe populations, looks at how we estimate quantities, discusses how hypotheses are tested, and provides a brief overview of two major frameworks of statistical analysis.
  • Enhanced coverage of genomic perspectives throughout helps students to stay informed on this developing area of study.
  • Entirely reillustrated in a clean, contemporary style that enhances the content.